Recently widowed Benjamin "Ben" Mee (Matt Damon), still grieving his loss and dealing with the expulsion of his 14-year-old son, Dylan (Colin Ford), decides to make a fresh start by buying a new house. He tours many houses with his 7-year old daughter, Rosie (Maggie Elizabeth Jones), and his realtor, Mr. Stevens (J.B. Smoove), but finds none he likes until his daughter finds a listing with what seems like the perfect house. They drive to an old large house and Ben inspects the property, telling his realtor it is perfect. When they hear a lion roar, Stevens explains that the house comes with a zoo in the back, which closed several years before; if they want the house, they must buy the zoo.
Although reluctant at first, Mee decides to buy the zoo when he sees Rosie happily playing with the peacocks. Dylan, however, hates the idea of moving away from his friends and retreats into his artwork which has grown more macabre since the death of his mother. Ben's brother, Duncan (Thomas Haden Church), tries to dissuade him from the purchase, but Ben buys it anyway. The zoo staff, led by the 28-year old head keeper, Kelly Foster (Scarlett Johansson), help to start renovating the zoo with the intent to reopen it to the public. When Kelly confronts Ben about why he bought a zoo, knowing nothing about running one, Ben simply responds "Why not?". Dylan befriends Kelly's 13-year old homeschooled cousin, Lily Miska (Elle Fanning), who develops a romantic interest that Dylan is oblivious to.
A strict USDA inspector, Walt Ferris (John Michael Higgins), arrives for a surprise inspection and makes a list of repairs that would cost around $100,000, which Ben does not have. Rhonda Blair (Carla Gallo), the zoo's accountant, gossips that Ben will probably sell the zoo. The workers' morale sinks, fearing the property will be sold to a buyer that will close it down.
When Lily tells Dylan that she heard his family might be leaving, he is overjoyed, which hurts her feelings. Ben discovers that his wife bequeathed him an investment account, with instructions to use the money wisely while listening to his heart. Duncan advises him to walk away and start over with the money, but Ben decides to use the money to repair the zoo. While this lifts the zoo workers' morale, Dylan is unhappy about having to stay; he confronts his father, and a heated argument ensues. They reconcile the next morning and Dylan admits he misses Lily. Ben gives his son advice using his favorite principle, that you only need 20 seconds of courage to achieve great things. Ben realizes that instead of trying to start over by forgetting his wife, he should accept that she will always be a part of him.
Prior to the grand opening of the zoo, the facility passes a very stringent inspection from Ferris, who grudgingly wishes them good luck. Dylan, following his father's advice, confesses to Lily that he loves her and she forgives him. The evening prior to the opening, the worst rainstorm in 100 years is predicted to wash it out. The weather clears in time, but in the morning they are disappointed when no visitors arrive. They discover that a fallen tree has blocked the access road, with a large crowd of waiting visitors behind it. The staff help them to climb over the tree. There are so many people that they run out of tickets, forcing Ben and Kelly to look for more. They end up face to face in a shed, where Kelly admits to Ben that she is romantically interested in him but "can't get a handle on it". She kisses him, telling him that maybe they can do it again on New Year's Eve; Ben kisses her back, and tells her that he is looking forward to it.
Ben takes his children to the restaurant where he met their mother, explaining that it was the point where their existence became a possibility. He takes them through that day's encounter, where Ben worked up the nerve to talk to his future wife with "20 seconds of courage", and walks up to the table she was at. He visualizes her (Stephanie Szostak) sitting there, and asks her why such an amazing woman would talk to someone like him. She responds, "Why not?"
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