Percy recounts the story of Thalia and her great sacrifice in order to protect her friends. A young Annabeth, Luke, Grover and Thalia are running to Camp Half-Blood while being pursued by monsters. Thalia sacrifices herself to get the others into the camp and her father Zeus finds another way for her to live by transforming her into a pine tree, creating a magical border around the camp so that no other half-blood would suffer from a fate as she did. Back at the camp, Percy has started to feel like a "One Quest Wonder". Soon after, Chiron and Mr. D investigate a Cyclops that has somehow passed through the border, and the Cyclops is revealed to be Tyson, Percy's half-brother. After the campers pick on Percy and Tyson, the camp is suddenly attacked by a Colchis Bull, who breaks through the border and ravages Camp Half-Blood. Percy finally defeats it with the help of Tyson, Annabeth, and Clarisse. Following this, Luke Castellan (who had survived) suddenly appears and confronts Percy, trying to convert the son of Poseidon to his cause. When Percy refuses, Luke disappears.
The campers soon realize that Thalia's tree has been poisoned by Luke and that they have been left vulnerable to attacks. Percy asks Chiron about a prophecy he is involved in (for Luke had tried to bait him with the knowledge earlier), and subsequently visits the Oracle, who tells him of a prophecy of a half-blood of the eldest gods either saving or destroying Olympus. Percy tells Chiron about the prophecy, and Chiron then tells him that he is the only living half-blood of the eldest gods, so the half-blood in the Prophecy may refer to Percy himself. Annabeth and Grover then find a way to save the tree as they learn about the Golden Fleece which has the power to heal almost anything and they then propose a quest to Mr. D, in which they retrieve the Fleece from Polyphemus' Lair in Circeland in the Sea of Monsters and use it to heal Thalia's tree. Mr. D then announces the quest, but chooses Clarisse to lead the quest instead, much to Annabeth's and Percy's dismay. Percy then convinces Grover and Annabeth to accompany him on the quest, and Tyson joins them against Annabeth's wishes. Annabeth hails the Chariot of Damnation and Percy threatens the three drivers (the Graeae) to tell him of his prophecy, causing them to give him a set of coordinates in the Sea of Monsters before ejecting the group from the cab in Washington D.C. While walking down the street, Grover is kidnapped by Chris Rodriguez and taken to Luke. Meanwhile, Percy, Annabeth, and Tyson meet with Hermes and he gives them a thermos that releases the winds from the four corners of the earth, as well as a magical box sealer that, when taped around as a border, can make anything disappear. The three ride a Hippokampos to Luke's yacht, The Andromeda, where they are captured and locked in the brig. Percy frees them with the box sealer and they escape after fighting through Luke's men and a Manticore, using the thermos to escape on the ship.
The group is consumed by Charybdis after Tyson loses the thermos, and they discover Clarisse in the monster's stomach. Percy and Clarisse work together to escape Charybdis's gut, and soon they arrive at Circeland, Polyphemus' lair. Percy finds Grover and the five escape Polyphemus, retrieving the fleece and trapping him in his cave. Afterwards, Luke arrives and demands the fleece from Percy, who refuses. Luke shoots a crossbow bolt at Percy but Tyson takes the bolt in the chest before falling to the water below, much to Percy's shock.
Luke begins reviving Kronos as Percy laments over the loss of Tyson. Annabeth then encourages Percy to take leadership. The team then escapes captivity and a fight ensues. Percy grapples with Luke over the Fleece, but Luke easily gains the upper hand. Luke is suddenly thrown away by Tyson, revealed to have survived his wound, due to the water healing it, as he is Poseidon's son. The group watches in horror as Kronos rises from the sarcophagus. Kronos consumes Luke and Grover, before battling Percy. Percy realizes that Riptide is the "cursed blade" of the prophecy, and slays Kronos with it, who regurgitates Grover and Luke, the latter landing in Polyphemus' Lair. Their victory is short lived as Annabeth is stabbed from behind by the Manticore, who is killed in turn by Clarisse and Grover. Annabeth dies in Percy's arms, but is resurrected by the Fleece. Percy then gives the fleece to Clarisse and they return to Camp Half-Blood. Clarisse places the fleece on Thalia's tree with Annabeth protecting the tree over night. The group returns the next day to find Thalia alive, as the fleece healed Thalia too much and returned her to human form.
Percy realizes that Thalia is a child of Zeus, and therefore is another possible child of the prophecy about either preserving or destroying Olympus. The film ends shows the sarcophagus with Kronos' remains in it glowing, presumably showing that Kronos is still alive somehow.
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