In 1938, in Egypt a team of archaeologists are digging into a tomb to look for a very valuable artifact. A young boy falls through a hole, leading his father and the others to what they came for - the tablet of Ahkmenrah. Several local men see the tablet and say that if the archaeologists remove it, then "the end will come".
In present day New York City, Larry Daley is overseeing a nighttime event at the Museum of Natural History with Dr. McPhee in attendance. Larry gathers the favorite exhibits, including Teddy Roosevelt, Attila the Hun, Sacagawea, Dexter the Monkey, and Rexy the T-Rex skeleton. Larry meets a new wax figure Neanderthal modeled after himself named Laaa. Ahkmenrah had earlier warned of the tablet's corrosion, causing the exhibits to act abnormally. The corrosion causes chaos, with the attendees having to flee from dangers caused by the affected exhibits. When Larry returns home, he catches his son Nicky throwing a party as the DJ. Nicky explains he doesn't intend to apply to college, wanting to take time to pursue being a DJ and to figure out what he wants to do with his life.
Larry researches the tablet and learns Cecil Fredericks, former security guard who tried stealing the tablet, was the boy present at its discovery. Larry finds Cecil in a retirement home along with his cohorts Gus and Reginald. Larry explains to Cecil what is happening and he recalls "the end will come" prophecy, realizing it was not an "end of the world" prophecy but a reference to an end to the magic. He suggests they consult Ahkmenrah's parents, who are in the British Museum. Larry later goes to Dr. McPhee, who is being fired, and convinces him to call the British Museum and allow Larry to travel there with Ahkmenrah and the tablet.
Larry and Nicky travel to London, meeting security guard Tilly, who lets them into the museum. When Larry gets into the museum, he sees that Laaa, Teddy, Sacagawea, Attila, Dexter, Jed, and Octavius have all stowed away to join on the adventure. The Tablet brings the British exhibits to life and the gang encounters a Triceratops skeleton which chases them. They are saved by Sir Lancelot. Unfortunately, Jed and Octavius fell into the ventilation chambers. Larry sends Dexter into the ventilation with Nicky's cell phone attached to his waist to find Jed and Octavius. Jed and Octavius land on a Pompeii exhibit just before it erupts. The gang fights off a nine-headed Xiangliu statue, which Larry defeats using a defibrillator. Meanwhile, Dexter urinates on the lava to save Jed and Octavius.
The gang finds the tombs of Ahkmenrah's parents, Merenkahre and Shepseheret. Merenkahre is at first hesitant to reveal the truth behind the Tablet, but he eventually admits it was a gift made for his son when he was a baby to keep the family together always. Endowed with the power of Khonsu, the tablet needs moonlight to retain its magic. Lancelot steals the tablet, mistaking it for the Holy Grail and then leaves to find Guinevere. The tablet continues to corrode, causing Teddy's hands to return to wax and Ahkmenrah to age. The gang splits up to find Lancelot, but Larry is caught by Tilly, who then locks Larry and Laaa in the security room. Larry starts to think about his relationship with Nicky; Laaa smashes the door's glass with his head to free them. The gang continue the search, but are cornered by the Trafalgar Square lion statues. Larry distracts them with a flashlight like in the cat video Jed and Octavius had previously watched on YouTube.
Lancelot has arrived at a local theater showing a Camelot production starring Hugh Jackman as King Arthur and Alice Eve as Guinevere; he runs onstage, trying to convince "Guinevere" to join him. They convince Lancelot he's mistaken, causing him to the roof. Larry and the gang attempt convincing him to adjust the tablet to receive moonlight. Larry convinces Lancelot to give him back the tablet to save his friends once the other exhibits all return to their lifeless state due to the tablet's corrosion. Larry then adjusts the tablet and it is restored by the moon, reanimating Teddy, Dexter, and the rest. Larry returns the tablet to Merenkahre after the New York exhibits explain that they believe the tablet and the pharaohs should stay together. On the way out, Larry catches Tilly and Laaa in a romantic moment. As they leave, Larry tells Tilly that, starting the next day, she will have the greatest job in the world. Back in New York, the exhibits accept their fate and say goodbye to Larry.
Three years later, a British Museum treasures exhibition event is happening at the American Museum of Natural History. Tilly shows up to McPhee's office with a box. McPhee explains that Larry left the museum and went on to become a teacher, after taking the blame for what happened when the exhibits went ballistic, granting McPhee his job back. Tilly opens the box and takes out the tablet. She brings Dr. McPhee to show him all the exhibits coming to life, including some new light displays. The revived exhibits then have a party. Outside, Larry pauses on the sidewalk to see the flashing lights from inside the museum; as he watches, he smiles.
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