In 1954, a United States Castle Bravo hydrogen bomb is used to destroy a giant creature surfacing from the ocean, covering it up as a nuclear bomb test. In 1999, Project Monarch scientists Ishiro Serizawa and Vivienne Graham go to the Philippines to investigate a colossal skeleton in a collapsed mine. Two egg-shaped spores are discovered, one broken open and leaving a trail to the sea. In Japan, the Janjira Nuclear Power Plant experiences unusual seismic activity. Joe Brody, the plant's supervisor, sends his wife, Sandra, and a team of technicians into the reactor to check the sensors. When the team is inside, the reactor is breached, releasing radioactive steam. Sandra and her team are unable to escape and the plant collapses into ruin.
Fifteen years later, Joe's son Ford is a US Navy explosive ordnance disposal officer, living in San Francisco with his wife Elle and son Sam. When Joe is arrested for trespassing in the Janjira quarantine zone, Ford returns to Japan. Convinced that the government covered up the true cause of the disaster, Joe convinces Ford to accompany him to their old home inside the zone to retrieve vital seismic data he recorded. They find the zone is not contaminated, but after recovering the data, soldiers appear and detain them in a Monarch facility within the plant's ruins. Inside, a giant winged creature emerges from a massive chrysalis and escapes, destroying the facility. Joe is injured and later dies. The media reports the incident as an earthquake.
A US Navy task force led by Admiral William Stentz on the aircraft carrier USS Saratoga starts a search for the creature, which they dub "MUTO" (Massive Unidentified Terrestrial Organism) and Serizawa, Graham and Ford go on-board to assist. The scientists reveal the 1954 appearance of Godzilla, a prehistoric alpha predator awoken by a deep sea expedition, how early nuclear tests were really attempts to kill it, that Project Monarch was formed secretly to study Godzilla and that the MUTO caused the Janjira plant's destruction. Ford reveals that Joe had monitored echolocation signals that indicated the MUTO was communicating with something.
A US Army Special Forces team finds the MUTO feeding off the wreckage of a Russian nuclear submarine it deposited in a forest near Honolulu. Ford waits at Honolulu airport for a flight back to San Francisco, when a battle breaks out between the military and the MUTO. Godzilla arrives, causing a tsunami that devastates Waikiki. After briefly fighting Godzilla, the MUTO flies away. Meanwhile, at the Yucca Mountain nuclear waste repository, the second found pod hatches into a larger, wingless female MUTO, which devastates Las Vegas. The scientists conclude that the two MUTOs were communicating and will meet to breed.
The task force follows Godzilla, projecting that the monsters will meet in San Francisco Bay. Stenz approves a plan to use a nuclear explosion to kill all three monsters, over the scientists' objections. Ford returns with the military to California and joins a team delivering warheads to San Francisco by train. After the female MUTO destroys the train, the one remaining warhead is airlifted to San Francisco and is armed. It's taken by the male MUTO and the female MUTO constructs a nest around it in the downtown.
The military fails to stop Godzilla when it arrives at the Golden Gate Bridge and pulls back to allow the monsters to fight. During the ensuing monsters' battle, a team of soldiers, including Ford, enter the city by HALO jump to attempt to disarm the warhead. When they fail, they take the warhead onto a boat for disposal at sea. Ford incinerates the nest, causing the female to leave the battle in a rage. Godzilla then uses its tail to smash the male MUTO into a building, killing him. The female MUTO wipes out the team on the boat, but before she can kill Ford, Godzilla kills her using its atomic breath. With the fight over, Godzilla collapses from exhaustion. Ford pilots the boat out to sea, and is rescued by a helicopter before the warhead detonates.
The next day, Ford is reunited with his family. The military cleans up the ruins while people look for other survivors. Godzilla, thought to be dead, awakens and returns to the Pacific Ocean.
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