A race of aliens called Mimics have taken over continental Europe. Major William Cage (Tom Cruise), a Public Affairs officer in the United States Marine Corps, seconded to the NATO-led United Defense Forces (UDF), is summoned to London to meet with General Brigham (Brendan Gleeson). Brigham states the UDF's intention to launch Operation Downfall against the Mimics. When he orders Cage to cover combat on the beaches of France, Cage objects to the dangerous assignment, and threatens to portray the General in an unfavorable light, for which he is arrested and knocked out. He awakens in handcuffs at a forward operating base at Heathrow Airport. Cage meets Master Sergeant Farell (Bill Paxton) and a squad of rejects known as J Squad. Brigham has sent a note to Farell, claiming that Cage is actually a private and a deserter who will falsely claim to be an officer. Farell assigns Cage to J Squad.
Cage is dropped in with the first wave, unaware that the Mimics have anticipated the invasion. Cage is attacked by an unusually large "Alpha" Mimic and uses a Claymore mine to destroy it. The Mimic's blood douses him after the mine explodes, and Cage wakes at Heathrow the previous morning. Realizing he now knows the outcome of the operation, Cage unsuccessfully tries to convince his squad mates that the invasion will fail and everyone will die. Each time he dies, he wakes up the previous morning. In one such loop he saves Sergeant Rita Vrataski (Emily Blunt), a famous soldier known as "Full Metal Bitch" and the "Angel of Verdun", in reference to the battle at Verdun where Vrataski slew hundreds of Mimics. She recognizes Cage's ability to anticipate events and tells him to locate her when he "wakes up".
Cage finds Vrataski and explains that they know each other from the future invasion. Vrataski confides in Cage that she too possessed the ability to reset, which allowed her to kill hundreds of Mimics at Verdun, but lost it after receiving a blood transfusion. Vrataski takes Cage to see Dr. Carter (Noah Taylor), a former government scientist and expert in Mimic biology. The doctor explains to Cage the time-loop situation, saying that the Mimics possessed the ability to reset time whenever an Alpha was killed, and by being covered in an Alpha's blood during the invasion, Cage had inherited the ability. Vrataski tells Cage that he will begin to experience "visions" of the aliens' hive mind called the Omega. Cage eventually sees the Omega in a Bavarian Alps dam. Through training with Vrataski, Cage's combat proficiency increases along with his memory gained from the previous loops.
In one loop, feeling the fight is pointless, Cage decides not to meet Vrataski and instead goes to a London pub. While there, civil defense sirens draw Cage out of the pub where he witnesses a massive attack on London. Cage now realizes the Mimics true plan: a large–scale amphibious assault on London while Operation Downfall forces are destroyed. After Cage is killed in London during the Mimic attack, Vrataski and Cage work on a plan to make it off the beach. Eventually, they make their way inland toward the dam.
After numerous loops end in disaster at a French farmhouse with Cage and Vrataski being repeatedly killed by Mimics, Vrataski convinces Cage that her life is expendable and he gives up on saving her. In the next loop, Cage goes alone to kill the Omega, abandoning Vrataski and the rest of the invasion to doom on the beach. When he arrives at the dam, he discovers that the Omega is not present, and is ambushed by an Alpha that tries to steal his blood and prevent him from resetting the day. Cage manages to kill himself by drowning underneath damaged water pipes. During the next loop, Cage tells Vrataski and Carter that his vision was a trick to ambush him and take away his ability to reset. Vrataski forces Carter to explain to Cage that during Carter's employment in Whitehall, General Brigham supervised the construction of a prototype transponder. Carter states the device can theoretically ascertain the location of the Omega if injected into an Alpha Mimic or someone with the reset ability. Vrataski and Cage infiltrate the Ministry of Defence and threaten to kill Brigham unless he gives them the transponder. Brigham relents after several failed loop attempts to convince him, and feigns co-operation. Upon walking out from the building, they are surrounded by armed soldiers who have been alerted to their presence by Brigham. In order to escape, Vrataski shoots Cage in the head to reset the day, and in their next loop, instead flee through the basement garage in a stolen car.
While in flight, Vrataski stabs the transponder into Cage's leg and he "sees" that the Omega is actually located under the glass pyramid outside the Louvre in Paris. After crashing and being captured by UDF soldiers, Cage wakes, tied to a bed, having had a blood transfusion and realizing he has lost the ability to reset the day. After Vrataski frees him, Cage convinces members from J Squad to help destroy the Omega. On their flight to Paris they are attacked by Mimics and crash-land. Surviving members of the squad sacrifice themselves to buy Vrataski and Cage time. Both make it beneath the Louvre, where they are hunted by an Alpha. Cage goes to kill the Omega as Vrataski distracts the Alpha and is killed. The Alpha then catches up to and mortally wounds Cage, but not before he primes and drops a grenade belt into the mouth of the Omega, destroying it and, as a result, all other Mimics.
As the Omega disintegrates, Cage's body absorbs its blood and he wakes up en route to his initial meeting with Brigham the day before. He attends a press conference where Brigham announces that a power surge was detected in Paris after which all Mimic activity ceased. Brigham says that the Mimics have been defeated, and the Russian and Chinese armies are advancing into Europe without resistance. Cage is never arrested and visits Vrataski at the Heathrow base. She asks what he wants, as always when not recognizing him, then Cage smiles and laughs.
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