In 2005, scientists discover an extrasolar planet, which they name Planet-G, with conditions similar to Earth. In 2006 NASA transmits a powerful signal from a communications array in Hawaii. Meanwhile, slacker Alex Hopper gets arrested while attempting to impress Sam Shane, daughter of U.S. Pacific Fleet commander Admiral Terrance Shane. Stone Hopper, Alex's older brother and a naval officer, is infuriated at Alex's lack of motivation and forces Alex to join him in the U.S. Navy.
By the present day in the year 2012, Alex is a hothead lieutenant and tactical action officer aboard the Arleigh Burke-class destroyer USS John Paul Jones, while Stone is a commander and the commanding officer of USS Sampson. Alex is in a relationship with Sam and is in danger of being discharged from the Navy. Their ships join the 2012 Rim of the Pacific Exercise (RIMPAC) in Hawaii.
During the exercises five alien spacecraft arrive in response to the NASA signal. Their communications ship collides with a satellite and crashes in Hong Kong, causing heavy damage and casualties, while the four others land in the water near Hawaii. Sampson, John Paul Jones, and the JMSDF Kongo class destroyer Myoko investigate. Alex, Raikes, and Beast are sent to investigate, but when Alex approaches, the alien mothership reacts and creates a force field that traps the three warships. The Navy issues a blast from a fog horn, which is answered in kind by a sonic blast from one of the alien ships. The Navy opens fire on the alien fleet, which then reevaluates the Earth ships' friend or foe status, changing the monitor color for the Navy ships from green to red. The aliens destroy the Sampson and Myoko, and damage John Paul Jones, killing the commanding and executive officers. Alex sees his brother die and returns to his ship, where he is forced to take command, albeit reluctantly and to the disbelief of the crew, as the most senior officer left on the ship. Enraged, Alex orders an attack, but Beast manages to convince him to recover the survivors from Myoko, including Captain Nagato, with whom Alex is in a rivalry. The alien ships sends weapons to the islands that destroy the military equipment and island infrastructure, killing a large number of innocent civilians when they destroy the freeway leading to a military base.
In Oahu, Sam, a physical therapist, is accompanying retired U.S. Army veteran and double amputee Mick Canales, on a hike to help him adapt to his prosthetic legs. At the sea, they capture an humanoid alien who is seemingly unconscious. The alien attacks Alex when startled by a bright light and establishes a mind meld with him, showing him how the aliens have attacked and conquered other planets. Another alien infiltrates the ship, but the crew manages to blast him off using a 5"/54 caliber Mark 45 gun. Sam and Mick run into some police officers who order them off the mountain. The cops head up the mountain and are ambushed and killed by aliens. Later, Sam and Mick run into scientist Cal Zapata, who informs them that the aliens have killed his grad student and are using the communications array to signal back to their home planet.
Back on the John Paul Jones and thanks to the briefly captured alien, the humans learn that the aliens are sensitive to sunlight. However, they face difficulty because the alien force field has disabled radar functions. Nagata suggests using NOAA's tsunami warning buoys around Hawaii to track the aliens as he has been using it to track other countries ships for every RIMPAC game. Mick forces Zapata to retrieve a spectrum analyser that allows Sam and Mick to contact the John Paul Jones and relay that the communications satellite will be on position in four hours and the aliens will be able to contact their planet. Zapata manages to take the analyser, and he is caught by an alien but escapes. During a night battle, the John Paul Jones sinks two alien ships but is unable to hit the third. Alex decides to lure the third alien vessel close to shore at dawn, where he and Nagata shoot out its bridge windows as the sunrise blinds the aliens. John Paul Jones destroys the last alien escort vessel only to fall prey to their mothership's long range weapons.
Realizing the aliens are attempting to contact a larger invasion fleet, Alex and the survivors of the John Paul Jones and Myoko assume command of USS Missouri, a decommissioned battleship turned into a museum ship. They reactivate the battleship with the aid of the retired veterans preserving her, but they end up coming face to face with the alien mothership. In the ensuing battle, the Missouri severely damages the alien mothership, destroying its force field in process. Alex uses the last shell to destroy the communications array on the island, leaving the Missouri defenseless. With the force field down, Admiral Shane scrambles the fleet and F-18 fighters from the RIMPAC fleet, who arrive and save the crew by eliminating the alien threat.
A ceremony is held to honor the Navy personnel, where Alex is promoted to lieutenant commander. He is presented with a Silver Star and also accepts Stone's posthumous Navy Cross. After the ceremony, Alex is given an offer to become a Navy SEAL, and then asks Admiral Shane for his daughter's hand in marriage. The admiral initially refuses but invites Alex to lunch to discuss the matter, referencing how Alex and Sam met.
In the post-credits, three schoolchildren and a handyman in Scotland are running off in terror as an alien hand reaches out of an object they have just opened.
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